Okay. So his meal was stolen, he is literally begging to have some of it back, and it's adorable to watch. Appreciate the full explanation of behavior, that's awesome, but you just described why wolves become beggars.
That wolf is not begging. It's running a cost-benefit calculation that wildlife biologists have been studying for 30 years.
What you're watching is kleptoparasitism. The wolf likely made this kill, and the grizzly walked up and took it. In Yellowstone, grizzlies usurp wolf kills constantly, sometimes within hours. One Yellowstone biologist watched a single bear hold 24 wolves off a bull elk they had brought down. The wolves sat and waited their turn.
The "playful puppy" posture is the wolf reading the odds. A hunt is dangerous and usually fails. Staying near a carcass a bear has claimed costs almost nothing. So the wolf lowers its body, keeps its distance, and waits for the bear to eat its fill and leave scraps. That submissive crouch is the optimal move, not affection.
The counterintuitive part: this theft makes wolves kill less, not more. For 20 years researchers assumed bears stealing carcasses forced wolves to hunt more often to make up the loss. Then a study across Yellowstone and Sweden found the opposite. Brown bear presence lowers the wolf kill rate in both ecosystems. Same finding, two continents, which is what convinced the biologists it was real.
Wolves put extra food on the landscape for bears. Bears give nothing back. They take a share of the shared prey and pirate the kills outright. For a Yellowstone boar, meat can run as high as 80 percent of the diet, and a lot of that protein arrives pre-hunted by a pack that did the dangerous work.
What looks like a puppy asking for a treat is one apex predator subsidizing another, and getting nothing in return.